Afterbreakingoutofarehabilitationzoo,Frankie,asquirrelwithalltheangles,returnstohisforest.Helearnsfromhisdim-wittedbestfriend,Cody,aporcupineandstreet-con,thatwhilehewasinlock-up,theentireforestwaspickedcleanbyTheA.C.O.R.N.S.Institute(AlternativeCosmeticOilsforRegenerativeNaturalSkin-care).Noacornwasleftbehind.Withwinterjustaroundthecorner,itsapotentiallydevastatingblowtoallforestcritters.So,Frankiehasnochoicebuttobustintothebrandnew,state-of-the-art,rodent-proofA.C.O.R.N.S.ProcessingPlantandtakebackalloftheacorns!Thoughaseeminglyimpossibleheist,successwouldnotonlysavethelivesofhisforestfriends,butitwouldalsogiveFrankieachancetowinbacktheloveofhislife,Lola,alocalshowsquirrel.Sadly,LolahasbeenburntbytoomanyofFrankiesfalsepromisesinthepast.WiththesupportofFrankiesoldmentor,Mr.Bellwood,awealthybadger,FrankieandCodyassembleateamoftopnotchcrackersfromallovertheforest,including:abatnamedFlyBoy,atopnotchmunitionsexpert;Raitch,acomputerhackingalbinosnake,whoneverallowsherlackofappendagestogetintheway;Edsyatinyfamilyfrog,whoseabilitytofreezehimselfsolidfordaysonendwithoutharmisinvaluable;agermophobicFrenchratnamedLiam,whocancollapsehisskeletontogetintoanyspace;Chinesefather-daughterpangolinsnamedPangoandLin,thebestdiggersintheforest;andBellwoodshedgehogbutler,Oscarwhoknowshowtogethishandsonanythingthecrewneeds.Withwinterknockingattheirdoor,Frankieandhiscrewhaveonlythreedaystoprepareandexecutethisunprecedentedheist.Whentheylearnthattheprocessingoftheirforestsacornshasbeenpushedupaday,LolasrefusaltoparticipateleavesFrankieandhiscrewonecrittershort.Witheverythingontheline,FrankieandhiscrewofAnimalCrackerzsetouttopulloffthebiggestheistinanimalhistory!同(tóng )行的还有，项目(🆕)赞助人冯•杜姆（朱利(⏸)安•麦(🎖)克马洪JulianMcMahon饰）、宇航员本•格林姆(🥠)（麦克•(🍡)切克里斯MichaelChiklis饰）、博士的(🛸)前女(😻)友(yǒu )苏•斯通（杰西卡•阿尔(🈷)芭JessicaAlba饰(📒)）以及(👔)她(🉑)弟弟(dì )约翰尼(ní )•斯通（克里(😚)斯•埃文(🅾)斯ChrisEvans饰(shì )）。身后道(🥌)路不平(🕓)，他在后退的(🤛)时候(hòu )踩(👤)到(dào )一块石头(tóu )上，一个趔趄(jū )，忽然就(jiù )重重(chóng )摔倒了地上，手里拎(🏮)着的饭盒也(🥦)摔了出去，撒了一地。在婢女益(🤱)春促使下，两(liǎng )人终成(🍉)连理，并相偕(💔)私奔。街(jiē(💄) )旁的路灯(📈)隐匿在高大的树荫之(❗)中，光线昏暗，只有(yǒu )路上来来往往的车灯，间或能照(zhào )亮容恒的(💍)脸(🚏)。已经四十岁(🌐)的她不仅将自己的私人生(🌨)(shēng )活打理的井井有(⛺)条，店内的(de )大(🔪)小(🏎)事务亦(🚘)手到(🏊)擒(💃)来不在话下，但(🤜)唯独感情这一块依旧是空缺的状态，这究竟(🌐)是为什么(me )呢(📺)？花田圣矢...。平静明(😓)媚(🍭)的一(🏫)天，大(dà )海上(shàng )突然传来路(🅾)飞(fēi )（田中真弓配音）撕(🤯)心(🚃)裂肺的(de )哀(⏹)(āi )号，原来在他(🈹)睡(😞)觉的时候，他最最钟(📸)爱的(🏻)那顶草帽竟然(🔍)不(bú )翼而飞。若真有万一，他也(😚)只能(néng )亲自介入。刘超(🅾)饰(shì )演(👰)(yǎ(📶)n )马远征，前海军陆(🌁)战队连(lián )长(zhǎng )